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The get_(local|foreign)_related_value methods of GenericRelation must be reversed because it defines (from|to)_fields and associated related_fields in the reversed order as it's effectively a reverse GenericForeignKey itself. The related value methods must also account for the fact that referenced primary key values might be stored as a string on the model defining the GenericForeignKey but as integer on the model defining the GenericRelation. This is achieved by calling the to_python method of the involved content type in get_foreign_related_value just like GenericRelatedObjectManager does. Lastly reverse many-to-one manager's prefetch_related_querysets should use set_cached_value instead of direct attribute assignment as direct assignment might are disallowed on ReverseManyToOneDescriptor descriptors. This is likely something that was missed in f5233dc (refs #32511) when the is_cached guard was added. Thanks 1xinghuan for the report. Backport of 198b30168d4e94af42e0dc7967bd3259b5c5790b from main.