diff --git a/django/contrib/admin/static/admin/css/changelists.css b/django/contrib/admin/static/admin/css/changelists.css
index ae90943e93..141b79040d 100644
--- a/django/contrib/admin/static/admin/css/changelists.css
+++ b/django/contrib/admin/static/admin/css/changelists.css
@@ -1,14 +1,22 @@
 /* CHANGELISTS */
 
-#changelist {
+#changelist .changelist-form-container {
     display: flex;
+    flex-wrap: wrap;
     align-items: flex-start;
-    justify-content: space-between;
+    width: 100%;
 }
 
-#changelist .changelist-form-container {
+#changelist .changelist-form-container > div {
     flex: 1 1 auto;
-    min-width: 0;
+}
+
+#changelist .changelist-form-container:not(:has(#changelist-filter)) > div {
+    width: 100%;
+}
+
+#changelist .changelist-form-container:has(#changelist-filter) > div {
+    max-width: calc(100% - 270px);
 }
 
 #changelist table {
diff --git a/django/contrib/admin/static/admin/css/responsive.css b/django/contrib/admin/static/admin/css/responsive.css
index 9aa895316c..bf6cd6a56c 100644
--- a/django/contrib/admin/static/admin/css/responsive.css
+++ b/django/contrib/admin/static/admin/css/responsive.css
@@ -409,11 +409,15 @@ input[type="submit"], button {
 
     /* Changelist */
 
-    #changelist {
-        align-items: stretch;
+    #changelist .changelist-form-container {
         flex-direction: column;
     }
 
+    #changelist .changelist-form-container:has(#changelist-filter) > div {
+        max-width: 100%;
+        width: 100%;
+    }
+
     #toolbar {
         padding: 10px;
     }
@@ -436,8 +440,7 @@ input[type="submit"], button {
     }
 
     #changelist-filter {
-        position: static;
-        width: auto;
+        width: 100%;
         margin-top: 30px;
     }
 
diff --git a/django/contrib/admin/templates/admin/change_list.html b/django/contrib/admin/templates/admin/change_list.html
index 3b3ea408d3..b12f8ec583 100644
--- a/django/contrib/admin/templates/admin/change_list.html
+++ b/django/contrib/admin/templates/admin/change_list.html
@@ -56,29 +56,8 @@
     {% endif %}
     <div class="module{% if cl.has_filters %} filtered{% endif %}" id="changelist">
       <div class="changelist-form-container">
-        {% block search %}{% search_form cl %}{% endblock %}
-        {% block date_hierarchy %}{% if cl.date_hierarchy %}{% date_hierarchy cl %}{% endif %}{% endblock %}
-
-        <form id="changelist-form" method="post"{% if cl.formset and cl.formset.is_multipart %} enctype="multipart/form-data"{% endif %} novalidate>{% csrf_token %}
-        {% if cl.formset %}
-          <div>{{ cl.formset.management_form }}</div>
-        {% endif %}
-
-        {% block result_list %}
-          {% if action_form and actions_on_top and cl.show_admin_actions %}{% admin_actions %}{% endif %}
-          {% result_list cl %}
-          {% if action_form and actions_on_bottom and cl.show_admin_actions %}{% admin_actions %}{% endif %}
-        {% endblock %}
-        {% block pagination %}
-        <div class="changelist-footer">
-        {% pagination cl %}
-        {% if cl.formset and cl.result_count %}<input type="submit" name="_save" class="default" value="{% translate 'Save' %}">{% endif %}
-        {% endblock %}
-        </div>
-        </form>
-      </div>
-      {% block filters %}
-        {% if cl.has_filters %}
+        {% block filters %}
+          {% if cl.has_filters %}
           <search id="changelist-filter" aria-labelledby="changelist-filter-header">
             <h2 id="changelist-filter-header">{% translate 'Filter' %}</h2>
             {% if cl.is_facets_optional or cl.has_active_filters %}<div id="changelist-filter-extra-actions">
@@ -92,8 +71,31 @@
             </div>{% endif %}
             {% for spec in cl.filter_specs %}{% admin_list_filter cl spec %}{% endfor %}
           </search>
-        {% endif %}
-      {% endblock %}
+          {% endif %}
+        {% endblock %}
+        <div>
+          {% block search %}{% search_form cl %}{% endblock %}
+          {% block date_hierarchy %}{% if cl.date_hierarchy %}{% date_hierarchy cl %}{% endif %}{% endblock %}
+
+          <form id="changelist-form" method="post"{% if cl.formset and cl.formset.is_multipart %} enctype="multipart/form-data"{% endif %} novalidate>{% csrf_token %}
+          {% if cl.formset %}
+            <div>{{ cl.formset.management_form }}</div>
+          {% endif %}
+
+          {% block result_list %}
+            {% if action_form and actions_on_top and cl.show_admin_actions %}{% admin_actions %}{% endif %}
+            {% result_list cl %}
+            {% if action_form and actions_on_bottom and cl.show_admin_actions %}{% admin_actions %}{% endif %}
+          {% endblock %}
+          {% block pagination %}
+          <div class="changelist-footer">
+          {% pagination cl %}
+          {% if cl.formset and cl.result_count %}<input type="submit" name="_save" class="default" value="{% translate 'Save' %}">{% endif %}
+          {% endblock %}
+          </div>
+          </form>
+        </div>
+      </div>
     </div>
   </div>
 {% endblock %}
diff --git a/tests/admin_views/admin.py b/tests/admin_views/admin.py
index 95d16a4770..b6d009b388 100644
--- a/tests/admin_views/admin.py
+++ b/tests/admin_views/admin.py
@@ -431,6 +431,8 @@ class PodcastAdmin(admin.ModelAdmin):
     list_display = ("name", "release_date")
     list_editable = ("release_date",)
     date_hierarchy = "release_date"
+    list_filter = ("name",)
+    search_fields = ("name",)
     ordering = ("name",)
 
 
diff --git a/tests/admin_views/test_skip_link_to_content.py b/tests/admin_views/test_skip_link_to_content.py
index 3284b76495..5cec63307b 100644
--- a/tests/admin_views/test_skip_link_to_content.py
+++ b/tests/admin_views/test_skip_link_to_content.py
@@ -3,6 +3,8 @@ from django.contrib.auth.models import User
 from django.test import override_settings
 from django.urls import reverse
 
+from .models import Podcast
+
 
 @override_settings(ROOT_URLCONF="admin_views.urls")
 class SeleniumTests(AdminSeleniumTestCase):
@@ -125,3 +127,45 @@ class SeleniumTests(AdminSeleniumTestCase):
             )
             self.assertTrue(is_vertical_scrolleable)
             self.assertFalse(is_horizontal_scrolleable)
+
+    def test_skip_link_keyboard_navigation_in_changelist(self):
+        from selenium.webdriver.common.by import By
+        from selenium.webdriver.common.keys import Keys
+
+        Podcast.objects.create(name="apple", release_date="2000-09-19")
+        self.admin_login(
+            username="super", password="secret", login_url=reverse("admin:index")
+        )
+        self.selenium.get(
+            self.live_server_url + reverse("admin:admin_views_podcast_changelist")
+        )
+        selectors = [
+            "ul.object-tools",  # object_tools.
+            "search#changelist-filter",  # list_filter.
+            "form#changelist-search",  # search_fields.
+            "nav.toplinks",  # date_hierarchy.
+            "form#changelist-form div.actions",  # action.
+            "table#result_list",  # table.
+            "div.changelist-footer",  # footer.
+        ]
+        content = self.selenium.find_element(By.ID, "content-start")
+        content.send_keys(Keys.TAB)
+
+        for selector in selectors:
+            with self.subTest(selector=selector):
+                # Currently focused element.
+                focused_element = self.selenium.switch_to.active_element
+                expected_element = self.selenium.find_element(By.CSS_SELECTOR, selector)
+                element_points = self.selenium.find_elements(
+                    By.CSS_SELECTOR,
+                    f"{selector} a, {selector} input, {selector} button",
+                )
+                self.assertIn(
+                    focused_element.get_attribute("outerHTML"),
+                    expected_element.get_attribute("innerHTML"),
+                )
+                # Move to the next container element via TAB.
+                for point in element_points[::-1]:
+                    if point.is_displayed():
+                        point.send_keys(Keys.TAB)
+                        break